Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Pteropodidae Gray, 1821 - Old World fruit bats
         Genus Pteropus Brisson, 1762 - flying foxes
           Pteropus mariannus species group
             1 ‡Pteropus loochoensis Gray, 1870
             2 Pteropus mariannus Desmarest, 1822 - Mariana flying fox
             3 Pteropus pelewensis Andersen, 1908
             4 Pteropus tonganus Quoy & Gaimard, 1830 - insular flying fox
             5 Pteropus ualanus Peters, 1883
             6 Pteropus yapensis Andersen, 1908