Class Mammalia Linnaeus, 1758 - mammals
   Subclass Theria Parker & Haswell, 1897
     Order Chiroptera Blumenbach, 1779 - bats
       Family Pteropodidae Gray, 1821 - Old World fruit bats
         Genus Pteropus Brisson, 1762 - flying foxes
           Pteropus subniger species group
             01 Pteropus admiralitatum Thomas, 1894 - Admirality flying fox
             02 ‡Pteropus brunneus Dobson, 1878 - Percy Island flying fox
             03 Pteropus dasymallus Temminck, 1825 - Ryukyu flying fox
             04 Pteropus faunulus Miller, 1902
             05 Pteropus griseus (Geoffroy, 1810) - grey flying fox
             06 Pteropus howensis Troughton, 1931
             07 Pteropus hypomelanus Temminck, 1853 - small flying fox
             08 Pteropus ornatus Gray, 1870
             09 Pteropus pumilus Miller, 1911 - little golden-mantled fruit bat
             10 Pteropus speciosus Andersen, 1908
             11 ‡Pteropus subniger (Kerr, 1792) - lesser Mascarene flying fox