Division Opisthokonta Cavalier-Smith, 1987 see Clade Opisthokonta Cavalier-Smith 1987, nom. cl. inval.
   Superkingdom Holomycota Liu, Steenkamp, Brinkmann, Forget, Philippe & Lang, 2009
     Kingdom Nucleariae Tedersoo, Sánchez-Ramírez, Kõljalg, Bahram, Döring, Schigel, May, Ryberg & Abarenkov, 2018
       1 Phylum Nuclearida Tedersoo, Sánchez-Ramírez, Kõljalg, Bahram, Döring, Schigel, May, Ryberg & Abarenkov, 2018 [monotypic]
       2 Phylum Fonticulida Tedersoo, Sánchez-Ramírez, Kõljalg, Bahram, Döring, Schigel, May, Ryberg & Abarenkov, 2018 [monotypic]