Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Deuterostomia Grobben, 1908
Phylum Chordata Bateson, 1885 - chordates
Subphylum Craniata auct.
Infraphylum Vertebrata Lamarck, 1801 - vertebrates
Clade Pan-Gnathostomata Downs 2020
Superclass Gnathostomataᵀ Gegenbaur, 1874 - jawed vertebrates
[crown] Clade Eugnathostomata
Clade Pan-Osteichthyes Rowe 2004 [Moore & Near 2020]
Clade Osteichthyes Huxley 1880 [Moore & Near 2020]
Clade Pan-Actinopterygii Rowe 2004 [Moore & Near 2020]
Clade Actinopterygiiᵀ Woodward 1891 [Moore & Near 2020]
Clade Pan-Actinopteri Moore & Near 2020
Class Actinopteri Cope, 1871
Subclass Neopterygii Regan, 1923
Infraclass Teleosteomorpha Arratia, 2001
Division Teleostei Müller, 1845
Supercohort Teleocephala de Pinna, 1996 - crown-group Teleostei
Cohort Clupeocephala Patterson & Rosen, 1977
Subcohort Otomorpha Wiley & Johnson, 2010
Superorder Ostariophysi Sagemehl, 1885
Section Otophysa Rosen & Greenwood, 1970
Superorder Cypriniphysae [monotypic]
Order Cypriniformesᵀ Bleeker, 1859 - carps, loaches, minnows, and relatives
Suborder Cyprinoideiᵀ Bleeker, 1859
Family Cyprinidaeᵀ Rafinesque, 1815 - carps
Subfamily Barbinae Bleeker, 1859 - barbels
Genus Barbusᵀ Cuvier & Cloquet, 1816
001 ?Barbus seymouri Tweddle & Skelton, 2008
002 †Barbus steinheimensis
003 †Barbus szchuanensis
004 Barbus ablabes (Bleeker, 1863)
005 Barbus aboinensis Boulenger, 1911
006 Barbus acuticeps Matthes, 1959
007 Barbus afrohamiltoni Crass, 1960
008 Barbus afrovernayi Nichols & Boulton, 1927
009 Barbus albanicus Steindachner, 1870
010 Barbus aliciae Bigorne & Lévêque, 1993
011 Barbus alluaudi Pellegrin, 1909
012 Barbus aloyi Roman, 1971
013 Barbus altidorsalis Boulenger, 1908
014 Barbus amanpoae Lambert, 1961
015 Barbus amatolicus Skelton, 1990
016 Barbus andrewi Barnard, 1937 - Cape whitefish
017 Barbus anema Boulenger, 1903
018 Barbus annectens Gilchrist & Thompson, 1917
019 Barbus anniae Lévêque, 1983
020 Barbus anoplus Weber, 1897
021 Barbus ansorgii Boulenger, 1904
022 Barbus apleurogramma Boulenger, 1911
023 Barbus arabicus Trewavas, 1941
024 Barbus arambourgi Pellegrin, 1935
025 Barbus arcislongae Keilhack, 1908
026 Barbus argenteus Günther, 1868
027 Barbus aspilus Boulenger, 1907
028 Barbus atakorensis Daget, 1957
029 Barbus atkinsoni Bailey, 1969
030 Barbus atromaculatus Nichols & Griscom, 1917
031 Barbus bagbwensis Norman, 1932
032 Barbus balcanicus Kotlík, Tsigenopoulos, Ráb & Berrebi, 2002
033 Barbus barbulus Heckel, 1847
034 Barbus barbusᵀ (Linnaeus, 1758) - barbel
035 Barbus barnardi Jubb, 1965
036 Barbus barotseensis Pellegrin, 1920
037 Barbus baudoni Boulenger, 1918
038 Barbus bawkuensis Hopson, 1965
039 Barbus bergi Chichkoff, 1935
040 Barbus bifrenatus Fowler, 1935
041 Barbus bigornei Lévêque, Teugels & Thys van den Audenaerde, 1988
042 Barbus boboi Schultz, 1942
043 Barbus borysthenicus Dybowski, 1862
044 Barbus bourdariei Pellegrin, 1928
045 Barbus brachygramma Boulenger, 1915
046 Barbus brazzai Pellegrin, 1901
047 Barbus breviceps Trewavas, 1936
048 Barbus brevidorsalis Boulenger, 1915
049 Barbus brevilateralis Poll, 1967
050 Barbus brevipinnis Jubb, 1966 - shortfin barb
051 Barbus brichardi Poll & Lambert, 1959
052 Barbus bynni (Forsskål, 1775)
053 Barbus cadenati Daget, 1962
054 Barbus calidus Barnard, 1938 - Clanwilliam redfin
055 Barbus callensis Valenciennes, 1842 - Algerian barb
056 Barbus callipterus Boulenger, 1907 - Clipper barb
057 Barbus camptacanthus (Bleeker, 1863) - African redfinned barb
058 Barbus candens Nichols & Griscom, 1917
059 Barbus caninus Bonaparte, 1839
060 Barbus carcharhinoides Stiassny, 1991
061 Barbus carens Boulenger, 1912
062 Barbus carottae (Bianco, 1998)
063 Barbus carpathicus Kotlík, Tsigenopoulos, Ráb & Berrebi, 2002
064 Barbus castrasibutum Fowler, 1936
065 Barbus catenarius Poll & Lambert, 1959
066 Barbus caudosignatus Poll, 1967
067 Barbus cercops Whitehead, 1960
068 Barbus chicapaensis Poll, 1967
069 Barbus chiumbeensis Pellegrin, 1936
070 Barbus chlorotaenia Boulenger, 1911
071 Barbus choloensis Norman, 1925
072 Barbus ciscaucasicus Kessler, 1877
073 Barbus citrinus Boulenger, 1920
074 Barbus claudinae De Vos & Thys van den Audenaerde, 1990
075 Barbus clauseni Thys van den Audenaerde, 1976
076 Barbus collarti Poll, 1945
077 Barbus condei Mahnert & Géry, 1982
078 Barbus cyclolepis Heckel, 1837
079 Barbus dartevellei Poll, 1945
080 Barbus deguidei Matthes, 1964
081 Barbus deserti Pellegrin, 1909
082 Barbus devosi Banyankimbona, Vreven & Snoeks, 2012
083 Barbus dialonensis Daget, 1962
084 Barbus diamouanganai Teugels & Mamonekene, 1992
085 Barbus ditinensis Daget, 1962
086 Barbus dorsolineatus Trewavas, 1936
087 Barbus eburneensis Poll, 1941
088 Barbus elephantis Boulenger, 1907
089 Barbus ensis Boulenger, 1910
090 Barbus ercisianus Karaman, 1971
091 Barbus erubescens Skelton, 1974 - Twee redfin
092 Barbus erythrozonus Poll & Lambert, 1959
093 Barbus ethiopicus Zolezzi, 1939
094 Barbus euboicus Stephanidis, 1950 - petropsaro
095 Barbus eurystomus Keilhack, 1908
096 Barbus eutaenia Boulenger, 1904
097 Barbus evansi Fowler, 1930
098 Barbus fasciolatus Günther, 1868 - African banded barb
099 Barbus fasolt Pappenheim, 1914
100 Barbus figuigensis Pellegrin, 1913
101 Barbus foutensis Lévêque, Teugels & Thys van den Audenaerde, 1988
102.1 Barbus fritschii Günther, 1874
102.2 Barbus harterti Günther, 1901
103 Barbus gananensis Vinciguerra, 1895
104 Barbus gestetneri Banister & Bailey, 1979
105 Barbus girardi Boulenger, 1910
106 Barbus goktschaicus Kessler, 1877
107 Barbus greenwoodi Poll, 1967
108 Barbus gruveli Pellegrin, 1911
109 Barbus grypus Heckel, 1843
110 Barbus guildi Loiselle, 1973
111 Barbus guineensis Pellegrin, 1913
112 Barbus guirali Thominot, 1886
113 Barbus gulielmi Boulenger, 1910
114 Barbus gurneyi Günther, 1868
115 Barbus haasi Mertens, 1925
116 Barbus haasianus David, 1936 - sickle barb
117 Barbus holotaenia Boulenger, 1904 - African longstripe barb
118 Barbus hospes Barnard, 1938 - Namaquab barb
119 Barbus huguenyi Bigorne & Lévêque, 1993
120 Barbus huloti Banister, 1976
121 Barbus hulstaerti Poll, 1945 - butterfly barb
122 Barbus humeralis Boulenger, 1902
123 Barbus humilis Boulenger, 1902
124 Barbus humphri Banister, 1976
125 Barbus inaequalis Lévêque, Teugels & Thys van den Audenaerde, 1988
126 Barbus innocens Pfeffer, 1896
127 Barbus issenensis Pellegrin, 1922
128 Barbus iturii Holly, 1929
129 Barbus jacksoni Günther, 1889
130 Barbus jae Boulenger, 1903 - Dja barb
131 Barbus janssensi Poll, 1976
132 Barbus jubbi Poll, 1967
133 Barbus kamolondoensis Poll, 1938
134 Barbus kerstenii Peters, 1868
135 Barbus kessleri (Steindachner, 1866)
136 Barbus kissiensis Daget, 1954
137 Barbus ksibi Boulenger, 1905
138 Barbus kuiluensis Pellegrin, 1930
139 Barbus lacerta Heckel, 1843
140 Barbus lagensis (Günther, 1868)
141 Barbus lamani Lönnberg & Rendahl, 1920
142 Barbus laticeps Pfeffer, 1889
143 Barbus lauzannei Lévêque & Paugy, 1982
144 Barbus leonensis Boulenger, 1915
145 Barbus lepineyi Pellegrin, 1939
146 Barbus leptopogon Schimper, 1834
147 Barbus liberiensis Steindachner, 1894
148 Barbus lineomaculatus Boulenger, 1903
149 Barbus longiceps Valenciennes, 1842
150 Barbus longifilis Pellegrin, 1935
151 Barbus lornae Ricardo-Bertram, 1943
152 Barbus lorteti Sauvage, 1882
153 Barbus loveridgii Boulenger, 1916
154 Barbus luapulae Fowler, 1958
155 Barbus lufukiensis Boulenger, 1917
156 Barbus luikae Ricardo, 1939
157 Barbus lujae Boulenger, 1913
158 Barbus lukindae Boulenger, 1915
159 Barbus lukusiensis David & Poll, 1937
160 Barbus luluae Fowler, 1930
161 Barbus macedonicus Karaman, 1928
162 Barbus machadoi Poll, 1967
163 Barbus macinensis Daget, 1954
164 Barbus macroceps Fowler, 1936
165 Barbus macrolepis Pfeffer, 1889
166 Barbus macrops Boulenger, 1911 - blackstripe barb
167 Barbus macrotaenia Worthington, 1933
168 Barbus magdalenae Boulenger, 1906
169 Barbus magniatlantis Pellegrin, 1919
170 Barbus manicensis Pellegrin, 1919
171 Barbus mariae Holly, 1929 - rhinofish
172 Barbus marmoratus David & Poll, 1937
173 Barbus martorelli Roman, 1971
174 Barbus massaensis Pellegrin, 1922
175 Barbus matthesi Poll & Gosse, 1963
176 Barbus mattozi Guimarães, 1884
177 Barbus mawambi Pappenheim, 1914
178 Barbus mawambiensis Steindachner, 1911
179 Barbus mediosquamatus Poll, 1967
180 Barbus melanotaenia Stiassny, 1991
181 Barbus meridionalis Risso, 1827
182 ‡Barbus microbarbis David & Poll, 1937
183 Barbus microterolepis Boulenger, 1902
184 Barbus mimus Boulenger, 1912
185.1 Barbus miolepis Boulenger, 1902
185.2 Barbus rubrostigma Poll & Lambert, 1964
186 Barbus mirabilis Pappenheim, 1914
187 Barbus mocoensis Trewavas, 1936
188 Barbus mohasicus Pappenheim, 1914
189 Barbus motebensis Steindachner, 1894 - Marico barb
190 Barbus moulouyensis Pellegrin, 1924
191 Barbus multilineatus Worthington, 1933
192 Barbus musumbi Boulenger, 1910
193 Barbus myersi Poll, 1939
194 Barbus nanningsi (de Beaufort, 1933)
195 Barbus nasus Günther, 1874
196 Barbus neefi Greenwood, 1962
197 Barbus neglectus Boulenger, 1903
198 Barbus neumayeri Fischer, 1884
199 Barbus nigeriensis Boulenger, 1903
200 Barbus nigrifilis Nichols, 1928
201 Barbus nigroluteus Pellegrin, 1930
202 Barbus niluferensis Turan, Kottelat & Ekmekçi, 2009
203 Barbus niokoloensis Daget, 1959
204 Barbus nounensis Van den Bergh & Teugels, 1998
205 Barbus nyanzae Whitehead, 1960
206 Barbus okae (Fowler, 1949)
207 Barbus oligogrammus David, 1937
208 Barbus oligolepis Battalgil, 1941
209 Barbus olivaceus Seegers, 1996
210 Barbus osseensis Nagelkerke & Sibbing, 2000
211 Barbus owenae Ricardo-Bertram, 1943
212 Barbus oxyrhynchus Pfeffer, 1889
213 Barbus pagenstecheri Fischer, 1884
214 Barbus pallaryi Pellegrin, 1919
215 Barbus pallidus Smith, 1841
216 Barbus paludinosus Peters, 1852
217 Barbus papilio Banister & Bailey, 1979
218 Barbus parablabes Daget, 1957
219 Barbus parajae Van den Bergh & Teugels, 1998
220 Barbus parawaldroni Lévêque, Thys van den Audenaerde & Traoré, 1987
221 Barbus paytonii Boulenger, 1911
222 Barbus pellegrini Poll, 1939
223 Barbus peloponnesius Valenciennes, 1842
224 Barbus perince Rüppell, 1835
225 Barbus petchkovskyi Poll, 1967
226 Barbus petenyi Heckel, 1852
227 Barbus petitjeani Daget, 1962
228 Barbus platyrhinus Boulenger, 1900
229 Barbus plebejus Bonaparte, 1839 - Italian barbel
230 Barbus pleurogramma Boulenger, 1902
231 Barbus pobeguini Pellegrin, 1911
232 Barbus poechii Steindachner, 1911
233 Barbus potamogalis (Cope, 1867)
234 Barbus prespensis Karaman, 1924 - briána
235 Barbus prionacanthus Mahnert & Géry, 1982
236 Barbus pseudotoppini Seegers, 1996
237 Barbus puellus Nichols & Boulton, 1927
238 Barbus pumilus Boulenger, 1901
239 Barbus punctitaeniatus Daget, 1954
240 Barbus pygmaeus Poll & Gosse, 1963
241 Barbus quadrilineatus David, 1937
242 Barbus quadripunctatus Pfeffer, 1896
243.1 Barbus radiatus Peters, 1853
243.2 Barbus profundus Greenwood, 1970
244 Barbus raimbaulti Daget, 1962
245 Barbus rebeli Koller, 1926
246 Barbus reinii Günther, 1874
247 Barbus rhinophorus Boulenger, 1910
248 Barbus rohani Pellegrin, 1921
249 Barbus rosae Boulenger, 1910
250 Barbus roussellei Ladiges & Voelker, 1961
251 Barbus rouxi Daget, 1962
252 Barbus ruasae Pappenheim, 1914
253 Barbus sacratus Daget, 1963
254 Barbus salessei Pellegrin, 1908
255 Barbus sensitivus Roberts, 2010
256 Barbus serengetiensis Farm, 2000
257 Barbus serra Peters, 1864 - sawfin
258 Barbus sexradiatus Boulenger, 1911
259 Barbus somereni Boulenger, 1911
260 Barbus sperchiensis Stephanidis, 1950
261 Barbus stanleyi Poll & Gosse, 1974
262 Barbus stappersii Boulenger, 1915
263 Barbus stauchi Daget, 1967
264 Barbus stigmasemion Fowler, 1936
265 Barbus stigmatopygus Boulenger, 1903
266 Barbus strumicae Karaman, 1955
267 Barbus subinensis Hopson, 1965
268 Barbus sublineatus Daget, 1954
269 Barbus subquincunciatus Günther, 1868
270 Barbus sylvaticus Loiselle & Welcomme, 1971
271 Barbus syntrechalepis (Fowler, 1949)
272 Barbus taeniopleura Boulenger, 1917
273 Barbus taeniurus Boulenger, 1903
274 Barbus tanapelagius Graaf, Dejen, Sibbing & Osse, 2000
275 Barbus tangandensis Jubb, 1954
276.1 Barbus tauricus Kessler, 1877
276.2 Barbus kubanicus Berg, 1912
277 Barbus tegulifer Fowler, 1936
278 Barbus tetrastigma Boulenger, 1913
279 Barbus teugelsi Bamba, Vreven & Snoeks, 2011
280 Barbus thamalakanensis Fowler, 1935
281 Barbus thessalus Stephanidis, 1971
282 Barbus thysi Trewavas, 1974
283 Barbus tiekoroi Lévêque, Teugels & Thys van den Audenaerde, 1987
284 Barbus tomiensis Fowler, 1936
285 Barbus tongaensis Rendahl, 1935
286 Barbus toppini Boulenger, 1916
287 Barbus trachypterus Boulenger, 1915
288 Barbus traorei Lévêque, Teugels & Thys van den Audenaerde, 1987
289 Barbus treurensis Groenewald, 1958 - Treur barb
290 Barbus trevelyani Günther, 1877 - border barb
291 Barbus trimaculatus Peters, 1852
292 Barbus trinotatus Fowler, 1936
293 Barbus trispiloides Lévêque, Teugels & Thys van den Audenaerde, 1987
294 Barbus trispilomimus Boulenger, 1907
295 Barbus trispilopleura Boulenger, 1902
296 Barbus trispilos (Bleeker, 1863)
297 Barbus tropidolepis Boulenger, 1900
298 Barbus turkanae Hopson & Hopson, 1982
299 Barbus tyberinus Bonaparte, 1839
300 Barbus unitaeniatus Günther, 1867 - redfinned barb
301 Barbus urostigma Boulenger, 1917
302 Barbus urotaenia Boulenger, 1913
303 Barbus usambarae Lönnberg, 1907
304 Barbus vanderysti Poll, 1945
305 Barbus venustus Bailey, 1980
306 Barbus viktorianus Lohberger, 1929
307 Barbus viviparus Weber, 1897
308 Barbus waleckii Rolik, 1970
309 Barbus walkeri Boulenger, 1904
310 Barbus wellmani Boulenger, 1911
311 Barbus wurtzi Pellegrin, 1908
312 Barbus yeiensis Johnsen, 1926
313 Barbus yongei Whitehead, 1960
314 Barbus zalbiensis Blache & Miton, 1960
315 Barbus zanzibaricus Peters, 1868