Arthropoda Siebold, 1848 - arthropods
   Insecta Linnaeus, 1758 - insects
     Pterygota Lang, 1888 - winged insects see Infraclass Pterygota Lang, 1888
       Endopterygota Sharp, 1899
         Lepidoptera Linnaeus, 1758 - butterflies, moths
           Heteroneura Tillyard, 1918 see Infraorder Heteroneura Tillyard, 1918
             Papilionoidea Latreille, 1802
               Nymphalidae Rafinesque, 1815 - brush-footed butterflies
                 Nymphalinae Rafinesque, 1815
                   II see Tribe Nymphalini Rafinesque, 1815
                     Vanessa Fabricius, 1807