Arthropoda Siebold, 1848 - arthropods
   Insecta Linnaeus, 1758 - insects
     Pterygota Lang, 1888 - winged insects see Infraclass Pterygota Lang, 1888
       Endopterygota Sharp, 1899
         Lepidoptera Linnaeus, 1758 - butterflies, moths
           Heteroneura Tillyard, 1918 see Infraorder Heteroneura Tillyard, 1918
             Papilionoidea Latreille, 1802
               Nymphalidae Rafinesque, 1815 - brush-footed butterflies
                 1 Argynninae P.A.J. Duponchel, [1835] see Tribe Argynnini Swainson, 1833
                 2 Anetinae see Subfamily Danainae Boisduval, 1833
                 3 Nymphalinae Rafinesque, 1815
                 4 Eurytelinae see Tribe Eurytelini
                 5 Limenitidinae Behr, 1864
                 6 Cyrestiinae see Subfamily Cyrestinae Guenée, 1865
                 6.0 Marpesiinae see Subfamily Cyrestinae Guenée, 1865
                 7 Apaturinae Boisduval, 1840
                 8 Charaxinae Guenée, 1865