Natura - nature
  
 actualia - actual entities
    
 Mundus Plinius - physical world
      
 naturalia - natural bodies
        
 Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
          
 Clade neomura Cavalier-Smith, 1987
            
 Domain Eukaryota Chatton, 1925 - eukaryotes
              
 Clade discaria Cavalier-Smith, 2021
                
 Clade dorsates Cavalier-Smith, 2021
                  
 Clade podiates Cavalier-Smith, 2012
                    
 Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
                      
 Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
                        
 Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
                          
 Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
                            
 Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
                              
 Kingdom Animalia Linnaeus, 1758 - animals
                                
 Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
                                  
 Subkingdom Bilateria Hatschek, 1888 - bilaterians
                                    
 Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
                                      
 Infrakingdom Protostomia Grobben, 1908
                                        
 Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
                                          
 Panarthropoda Nielsen, 1995 - panarthropods
                                            
 Tactopoda Budd, 2001
                                              
 Phylum Arthropoda Siebold, 1848 - arthropods
                                                
 Euarthropoda
                                                  
 Antennulata
                                                    
 Subphylum Mandibulata Snodgrass, 1938
                                                      
 Infraphylum Pancrustacea Zrzavý & Štys, 1997
                                                        
 Altocrustacea
                                                          
 Miracrustacea
                                                            
 Superclass Hexapoda Latreille, 1825
                                                              
 Cercophora
                                                                
 Class Insecta Linnaeus, 1758 - insects
                                                                  
 Subclass Dicondylia Haeckel, 1866
                                                                    
 Infraclass Pterygota Lang, 1888
                                                                      
 Metapterygota
                                                                        
 Neoptera Martynov, 1923
                                                                          
 Eumetabola
                                                                            
 Holometabola
                                                                              
 Superorder Hymenopterida
                                                                                
 Order Hymenopteraᵀ Linnaeus, 1758 - ants, bees, and wasps
                                                                                  
 Neohymenoptera
                                                                                    
 Euhymenoptera
                                                                                      
 Suborder Apocrita Gerstaecker, 1867
                                                                                        
 Aculeata Latreille, 1802
                                                                                          
 Superfamily Vespoidea Latreille, 1802
                                                                                            
 Family Vespidaeᵀ Latreille, 1802
                                                                                              
 Genus Ancistrocerus Wesmael, 1836
                                                                                              
 Genus Brachygastra Perty, 1833
                                                                                              
 Genus Cephalodynerus Parker, 1965
                                                                                              
 Genus Dolichodynerus Bohart, 1939
                                                                                              
 Genus Eumenes Latreille, 1802
                                                                                              
 Genus Euodynerus Dalla Torre, 1904
                                                                                              
 Genus Euparagia Cresson, 1879
                                                                                              
 Genus Gastrodynerus Bohart, 1984
                                                                                              
 Genus Hypalastoroides Saussure, 1856
                                                                                              
 Genus Leptochiloides Bohart, 1940
                                                                                              
 Genus Leptochilus Saussure, 1853
                                                                                              
 Genus Leucodynerus Bohart, 1982
                                                                                              
 Genus Maricopodynerus Viereck, 1908
                                                                                              
 Genus Microdynerus Thomson, 1874
                                                                                              
 Genus Mischocyttarus Saussure, 1853
                                                                                              
 Genus Monobia Saussure, 1852
                                                                                              
 Genus Montezumia Saussure, 1852
                                                                                              
 Genus Odynerus Latreille, 1803
                                                                                              
 Genus "Odynerus" auct.
                                                                                              
 Genus Pachodynerus Saussure, 1870
                                                                                              
 Genus Parancistrocerus Bequaert, 1925
                                                                                              
 Genus Paranortonia Bequaert, 1940
                                                                                              
 Genus Polistes auct.
                                                                                              
 Genus Polybia Lepeletier, 1836
                                                                                              
 Genus Pseudepipona Saussure, 1856
                                                                                              
 Genus Pseudodynerus Saussure, 1855
                                                                                              
 Genus Pseudomasaris Ashmead, 1902
                                                                                              
 Genus Pterocheilus Klug, 1805
                                                                                              
 Genus Smeringodynerus Snelling, 1975
                                                                                              
 Genus Stenodynerus Saussure, 1863
                                                                                              
 Genus Symmorphus Wesmael, 1836
                                                                                              
 Genus Zethus Fabricius, 1804
                                                                                              
 1 Subfamily Stenogastrinae
                                                                                              
 2 Subfamily Polistinae (Lepeletier de Saint-Fargeau, 1835)
                                                                                              
 3 Subfamily Vespinaeᵀ Latreille, 1802