Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Superorder Hymenopterida
Order Hymenopteraᵀ Linnaeus, 1758 - ants, bees, and wasps
Neohymenoptera
Euhymenoptera
Suborder Apocrita Gerstaecker, 1867
Aculeata Latreille, 1802
Superfamily Vespoidea Latreille, 1802
Family Mutillidae Latreille - velvet ants
Genus Acanthomutilla Nonveiller, 1995
Genus Acanthophotopsis Schuster, 1958
Genus Acrophotopsis Schuster, 1958
Genus Allotilla Schuster, 1949
Genus Allotropidia Nonveiller, 1996
Genus Amblotropidia Nonveiller, 1995
Genus Anomophotopsis Schuster, 1949
Genus Antennotilla Bischoff, 1921
Genus Apteromutilla Ashmead, 1903
Genus Arcuatotilla Nonveiller, 1998
Genus Areotilla Bischoff, 1920
Genus Arnoldtilla Nonveiller, 1996
Genus Artiotilla Invrea, 1950
Genus Ascetotilla Brothers, 1971
Genus Astomyrmex Schwartz
Genus Atillum André, 1903
Genus Aureotilla Bischoff, 1920
Genus Australotilla Lelej, 1983
Genus Barymutilla André, 1901
Genus Bethsmyrmilla Krombein & Lelej, 1999
Genus Bischoffitilla Lelej, 2002
Genus Bisulcotilla Bischoff, 1921
Genus Blakeius Ashmead, 1903
Genus Bothriomutilla Ashmead, 1899
Genus Brachymutilla André, 1901
Genus Brahmatilla Lelej
Genus Calomutilla Mickel, 1952
Genus Cameronilla Lelej, 2001
Genus Carinomyrmosa Lelej, 1981
Genus Carinotilla Nonveiller, 1973
Genus Cephalomutilla André, 1909
Genus Cephalotilla Bischoff, 1921
Genus Ceratophotopsis Schuster, 1949
Genus Ceratotilla Bischoff, 1921
Genus Chaetomutilla Nonveiller, 1978
Genus Chaetotilla Schuster, 1949
Genus Chilemutilla Cambra & Quintero
Genus Chilephotopsis Cambra & Quintero
Genus Chrestomutilla Brothers, 1971
Genus Chrysotilla Bischoff, 1920
Genus Clinotilla Arnold, 1956
Genus Cockerellidia Lelej & Krombein, 1999
Genus Corytilla Arnold, 1956
Genus Cretavus Sharov, 1957
Genus Crevatus Sharov
Genus Ctenoceraea Nonveiller, 1993
Genus Ctenotilla Bischoff, 1921
Genus Curvitropidia Nonveiller, 1995
Genus Cystomutilla André, 1896
Genus Darditilla Casal, 1965
Genus Dasylabris Radoszkowski, 1885
Genus Dasylabroides André, 1901
Genus Dasymutilla Ashmead, 1899
Genus Dentilla Lelej, 1980
Genus Dentotilla Nonveiller
Genus Dilophotopsis Schuster, 1958
Genus Dimorphomutilla Ashmead, 1903
Genus Dolichomutilla Ashmead, 1899
Genus Edrionotus Radoszkowski, 1885
Genus Eosmicromyrmilla Lelej & Krombein, 2001
Genus Eotrogaspidia Lelej, 1996
Genus Ephuamelia Casal, 1968
Genus Ephuchaya Casal, 1968
Genus Ephucilla Lelej, 1995
Genus Ephuseabra Casal, 1968
Genus Ephusuarezia Casal, 1968
Genus Ephuta Say, 1836
Genus Ephutomma Ashmead, 1899
Genus Ephutomorpha André, 1903
Genus Erimyrmosa Lelej, 1984
Genus Eurymutilla Ashmead, 1899
Genus Euspinolia Ashmead, 1903
Genus Glossotilla Bischoff, 1920
Genus Guincomutilla Suarez
Genus Guineomutilla Suárez
Genus Gurisita Casal, 1970
Genus Gynandrotilla Arnold, 1946
Genus Hadrotilla Bischoff, 1920
Genus Hildbrandetia Özdikmen
Genus Hindustanilla Lelej, 2001
Genus Hoplocrates Mickel, 1937
Genus Hoplognathoca Suárez, 1962
Genus Hoplomutilla Ashmead, 1899
Genus Horcomutilla Casal, 1962
Genus Indratilla Lelej, 1993
Genus Invreiella Suárez
Genus Jaxartilla Lelej, 1984
Genus Karlidia Lelej, 1999
Genus Karlissaidia Lelej
Genus Karunaratnea Lelej
Genus Krombeinella Pate, 1947
Genus Krombeinidia Lelej, 1996
Genus Kudakrumia Krombein, 1979
Genus Kurzenkotilla Lelej
Genus Labidomilla André, 1903
Genus Laminatilla Pitts
Genus Lehritilla Lelej
Genus Leiomyrmosa Wasbauer, 1974
Genus Leucospilomutilla Ashmead, 1903
Genus Limaytilla Casal, 1964
Genus Liomutilla André, 1907
Genus Liotilla Bischoff, 1920
Genus Lobotilla Bischoff, 1920
Genus Lomachaeta Mickel, 1936
Genus Lophomutilla Mickel, 1952
Genus Lophostigma Mickel, 1952
Genus Lophotilla Bischoff, 1920
Genus Lynchiatilla Casal, 1963
Genus Macromyrme Lelej, 1984
Genus Mickelia Suárez, 1966
Genus Mickelomyrme Lelej, 1995
Genus Mimecomutilla Ashmead, 1903
Genus Montanomutilla Nonveiller, 1978
Genus Morsyma Fox, 1899
Genus Myrmilloides André, 1903
Genus Myrmosula Bradley, 1917
Genus Myrmotilla Bischoff, 1920
Genus Nanomutilla André, 1901
Genus Nanomyrme Lelej, 1977
Genus Nanotopsis Schuster, 1949
Genus Nemka Lelej, 1985
Genus Neomutilla Reed, 1898
Genus Neotrogaspidia Lelej, 1996
Genus Nonveilleridia Lelej, 1996
Genus Nordeniella Lelej
Genus Nothomyrmosa Krombein, 1979
Genus Nuristanilla Lelej, 1980
Genus Odontomutilla Ashmead, 1899
Genus Odontomyrme Lelej, 1983
Genus Odontophotopsis Viereck, 1903
Genus Odontotilla Bischoff, 1920
Genus Omotilla Invrea, 1943
Genus Orientidia Lelej, 1996
Genus Orientilla Lelej, 1979
Genus Pagdenidia Lelej, 1996
Genus Paglianotilla Lelej
Genus Pappognatha Mickel, 1939
Genus Paramyrmosa Saussure, 1880
Genus Patquiatilla Casal
Genus Pertyella Mickel, 1952
Genus Petersenidia Lelej, 1992
Genus Photomorphus Viereck, 1903
Genus Physetopoda Schuster, 1949
Genus Platymyrmilla André, 1901
Genus Ponerotilla Brothers, 1994
Genus Promecidia Lelej, 1996
Genus Promecilla André, 1903
Genus Protomutilla Bischoff, 1916
Genus Protophotopsis Schuster, 1947
Genus Protrogaspidia Lelej, 1996
Genus Psammotherma Latreille
Genus Pseudocephalotilla Bischoff, 1921
Genus Pseudolophotilla Nonveiller & Cetkovic, 1995
Genus Pseudomethoca Ashmead, 1896
Genus Pseudomyrmosa Suárez
Genus Pseudophotopsis André, 1896
Genus Ptilomutilla André, 1905
Genus Pygomilla Hammer, 1955
Genus Radoszkowskitilla Lelej
Genus Rasnitsynitilla Lelej
Genus Reedomutilla Mickel, 1964
Genus Rhopalomutilla André, 1901
Genus Ronisia Costa
Genus Scaptodactyla Burmeister, 1874
Genus Schusterphotopsis Pitts, 2003
Genus Seabratilla Casal, 1963
Genus Serendibiella Lelej
Genus Seriatospidia Nonveiller & Cetkovic, 1996
Genus Sigilla Skorikov, 1927
Genus Sinotilla Lelej, 1995
Genus Skorikovia Ovtchinnikov, 2002
Genus Smicromutilla Mickel, 1964
Genus Smicromyrme Thomson, 1870
Genus Smicromyrmilla Suárez, 1965
Genus Somaliatilla Nonveiller, 1996
Genus Spilomutilla Ashmead, 1903
Genus Spinulomutilla Nonveiller, 1994
Genus Spinulotilla Bischoff, 1920
Genus Squamulotilla Bischoff, 1920
Genus Standfussidia Lelej
Genus Stenomutilla André, 1896
Genus Stethophotopsis Pitts, 2000
Genus Suareztilla Casal, 1968
Genus Sulcotilla Bischoff, 1921
Genus Sylvotilla Viette
Genus Taimyrmosa Lelej
Genus Taiwanomyrme Tsuneki, 1993
Genus Tallium André, 1903
Genus Timulla Ashmead, 1899
Genus Tobantilla Casal, 1965
Genus Traumatomutilla André, 1901
Genus Tricholabiodes Radoszkowski, 1885
Genus Trispilotilla Bischoff, 1920
Genus Trogaspidia Ashmead, 1899
Genus Tropidotilla Bischoff, 1920
Genus Tsunekimyrme Lelej, 1995
Genus Tuberocoxotilla Nonveiller, 1980
Genus Vanhartenidia Lelej & van Harten
Genus Vianatilla Casal, 1962
Genus Viereckia Ashmead, 1903
Genus Wallacidia Lelej & Brothers
Genus Xystromutilla André, 1905
Genus Yamanetilla Lelej
Genus Zavatilla Tsuneki, 1993
Genus Zeugomutilla Chen, 1957
1 Subfamily Myrmosinae
2 Subfamily Pseudophotopsidinae
3 Subfamily Ticoplinae
4 Subfamily Rhopalomutillinae
5 Subfamily Sphaeropthalminae Schuster, 1949
6 Subfamily Myrmillinae
7 Subfamily Mutillinaeᵀ Latreille, 1802