Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Adephaga Schellenberg, 1806
Family Dytiscidae Leach, 1815 - predaceous diving beetles
Genus Agabetes Crotch, 1873
Genus Agabinus Crotch, 1873
Genus Agaporomorphus Zimmermann, 1921
Genus Anodocheilus Babington, 1841
Genus Bidessonotus Régimbart, 1895
Genus Brachyvatus Zimmermann, 1919
Genus Carrhydrus Fall, 1922
Genus Celina Aubé, in Dejean & Boisduval, 1837-1838
Genus Coptotomus Say, 1834
Genus Derovatellus Sharp, 1881
Genus Desmopachria Babington, 1841
Genus Eretes Laporte, 1832
Genus Hoperius Fall, 1927
Genus Hydrotarsus Falkenström, 1938
Genus Hydrotrupes Sharp, 1882
Genus Laccodytes Regimbart, 1895
Genus Liodessus Guignot, 1939
Genus Lioporeus Guignot, 1950
Genus Matus Aube, 1838
Genus Megadytes Sharp, 1882
Genus Meladema Laporte, 1835
Genus Neobidessus Young, 1967
Genus Neoclypeodytes Young, 1967
Genus Neoscutopterus Balfour-Browne, 1945
Genus Pachydrus Sharp, 1882
Genus Rhithrodytes Bameul, 1989
1 Subfamily Hydroporinae Aubé, 1836
2 Subfamily Laccophilinae Gistel, 1856
3 Subfamily Colymbetinae Erichson, 1837
4 Subfamily Dytiscinaeᵀ (Leach, in Brewster, 1815)