Natura - nature
actualia - actual entities
Mundus Plinius - physical world
naturalia - natural bodies
Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
Clade neomura Cavalier-Smith, 1987
Domain Eukaryota Chatton, 1925 - eukaryotes
Clade discaria Cavalier-Smith, 2021
Clade dorsates Cavalier-Smith, 2021
Clade podiates Cavalier-Smith, 2012
Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
Superkingdom Holozoa Lang, O'Kelly, Nerad, Gray & Burger, 2002
Clade filozoa Shalchian-Tabrizi, Minge, Espelund, Orr, Ruden, Jakobsen & Cavalier-Smith, 2008
Kingdom Animalia Linnaeus, 1758 - animals
Clade Planulozoa Wallberg, Thollesson, Farris & Jondelius 2004
Subkingdom Bilateria Hatschek, 1888 - bilaterians
Clade Nephrozoa Jondelius, Ruiz-Trillo, Baguñà & Riutort 2002
Infrakingdom Protostomia Grobben, 1908
Superphylum Ecdysozoa Aguinaldo, Turbeville, Linford, Rivera, Garey, Raff & Lake, 1997
Panarthropoda Nielsen, 1995 - panarthropods
Tactopoda Budd, 2001
Phylum Arthropoda Siebold, 1848 - arthropods
Euarthropoda
Antennulata
Subphylum Mandibulata Snodgrass, 1938
Infraphylum Pancrustacea Zrzavý & Štys, 1997
Altocrustacea
Miracrustacea
Superclass Hexapoda Latreille, 1825
Cercophora
Class Insecta Linnaeus, 1758 - insects
Subclass Dicondylia Haeckel, 1866
Infraclass Pterygota Lang, 1888
Metapterygota
Neoptera Martynov, 1923
Eumetabola
Holometabola
Order Coleoptera Linnaeus, 1758 - beetles
Suborder Adephaga Schellenberg, 1806
Family Carabidae Latreille, 1802 - ground beetles
Genus Abaris Dejean, 1831
Genus Agonoderus
Genus Agra Fabricius, 1801
Genus Amblycheila Say, 1830
Genus Amblygnathus Dejean, 1829
Genus Amerinus Casey, 1884
Genus Ameroduvalius Valentine, 1952
Genus Amphasia Newman, 1838
Genus Anatrichis Leconte, [1854]
Genus Anchomenus Bonelli, 1810 ex Stephens, 1827
Genus Anchonoderus Reiche, 1843
Genus Anillaspis Casey, 1918
Genus Anillinus Casey, 1918
Genus Anillodes Jeannel, 1963
Genus Apenes LeConte, 1851
Genus Apristus de Chaudoir, 1846
Genus Ardistomis Putzeys, 1846
Genus Atelothrus Sharp, 1903
Genus Axinopalpus Le Conte, 1846
Genus Aztecarpalus Ball, 1970
Genus "Blemus" Dejean, 1821, nom. nud.
Genus Broscodera Lindroth, 1961
Genus Calleida Latreille & Dejean, 1824
Genus Calybe de Laporte, 1834
Genus Colpodes MacLeay, 1825
Genus Coptodera Dejean, 1825
Genus Cyclotrachelus Chaudoir, 1838
Genus Cylindronotum Putzeys, 1846
Genus Darlingtonea Valentine, 1952
Genus Dercylinus Chaudoir, 1882[1883]
Genus Diacheila Motschulsky, 1844
Genus Dicheirus Eschscholtz, in Mannerheim, 1843
Genus Diplochaetus de Chaudoir, 1871
Genus Diplocheila Brullé, 1834
Genus Diplous Motschulsky, 1850
Genus Discoderus Zimmermann, in Leconte, 1852
Genus Eccoptoptera de Chaudoir, 1878
Genus Elaphropus Motschulsky, 1839
Genus Elliptoleus Bates, 1882
Genus Eucaerus LeConte, 1853
Genus Eudalia Laporte, 1868[1867?]
Genus Euphorticus Horn, 1881
Genus Euproctinus Leng & Mutchler, 1927
Genus Euryderus LeConte, 1846
Genus Eurylychnus Bates, 1891
Genus Evolenes LeConte, 1853
Genus Feronia Latreille, 1817
Genus Galerita Fabricius, 1801
Genus Gehringia Darlington, 1933
Genus Geopinus LeConte, 1848
Genus Hadrocarabus Thomson, 1875
Genus Halocoryza Alluaud, 1919
Genus Hartonymus Casey, 1914
Genus Helluomorphoides Ball, 1951
Genus Horologion Valentine, 1932
Genus Infernophilus Larson, 1969
Genus Inna Putzeys, 1863
Genus Lachnocrepis LeConte, 1853
Genus Lachnophorus Dejean, 1831
Genus Leptotrachelus Latreille, 1829
Genus Lophoglossus LeConte, 1852
Genus Loxandrus LeConte, 1852
Genus Mecodema Blanchard, 1853
Genus Megacephala Latreille, 1802
Genus Megodonthus
Genus Metrius Eschscholtz, 1829
Genus Micranillodes Jeannel, 1963
Genus Micratopus Casey, 1914
Genus Micrixys LeConte, 1854
Genus Mioptachys Bates, 1882
Genus Morion Latreille, 1810
Genus Myas Dejean, 1828
Genus Neaphaenops Jeannel, 1920
Genus Nelsonites Valentine, 1952
Genus Nemotarsus LeConte, 1853
Genus Nomius Laporte, 1835
Genus Notiobia Perty, 1830
Genus Onota Chaudoir, 1872
Genus Oodinus Motschulsky, 1865
Genus Ophryogaster Chaudoir, 1878
Genus Opisthius Kirby, 1837
Genus Oxycrepis Reiche, 1843
Genus Oxydrepanus Putzeys, 1866
Genus Oxypselaphus Chaudoir, 1843
Genus Ozaena Olivier, [1812]
Genus Pachyteles Perty, 1830
Genus Paranchus Lindroth, 1974
Genus Paratachys Casey, 1918
Genus Pelmatellus Bates, 1882
Genus Pentagonica Schmidt-Goebel, 1846
Genus Pericompsus LeConte, 1852
Genus Philorhizus Hope, 1838
Genus Phloeoxena Chaudoir, 1869
Genus Phrypeus Casey, 1924
Genus Physea Brulle, 1834
Genus Piesmus LeConte, 1848
Genus Piosoma LeConte, 1848
Genus Platidiolus Chaudoir, 1878
Genus Platypatrobus Darlington, 1938
Genus Pogonistes Chaudoir, 1871
Genus Pogonodaptus Horn, 1881
Genus Polpochila Solier, 1849
Genus Polyderis Motschulsky, 1862
Genus Porotachys Netolitzky, 1914
Genus Promecognathus Chaudoir, 1846
Genus Pseudamara Lindroth, 1968
Genus Pseudanophthalmus Jeannel, 1920
Genus Pseudaptinus Laporte, 1834
Genus Pseudomorpha Kirby, 1825
Subfamily Pseudomorphinae (Hope, 1838)
Genus Psydrus LeConte, 1846
Genus Rhadine LeConte, 1846
Genus Selenophorus Dejean, 1829
Genus Semiardistomis Kult, 1950
Genus Sericoda Kirby, 1837
Genus Stenocrepis Chaudoir, 1857
Genus Stenomorphus Dejean, 1831
Genus Stereocerus Kirby, 1837
Genus Tanystoma Motschulsky, 1845
Genus Tecnophilus Chaudoir, 1877
Genus Tetragonoderus Dejean, 1829
Genus Tetraleucus Casey, 1920
Genus Thalassotrechus Van Dyke, 1918
Genus Thalpius LeConte, 1851
Genus Troglanillus Jeannel, 1963
Genus Xenotrechus Barr & Krekeler, 1967
Genus Xestonotus LeConte, 1853
Genus Zacotus LeConte, 1869
02 Subfamily Cicindelinae - tiger beetles
03 Subfamily Carabinaeᵀ Latreille, 1802
04 Subfamily Cychrinae
05 Subfamily Nebriinae (Laporte, 1834)
06 Subfamily Notiophilinae
07 Subfamily Omophroninae (Bonelli, 1810)
08 Subfamily Elaphrinae (Latreille, 1802)
09 Subfamily Loricerinae Bonelli, 1810
10 Subfamily Scaritinae Bonelli, 1810
11 Subfamily Broscinae Hope, 1838
12 Subfamily Trechinae Bonelli, 1810
13 Subfamily Bembidiinae
14 Subfamily Pogoninae
15 Subfamily Patrobinae (Kirby, 1837)
16 Subfamily Perigoninae
17 Subfamily Anchonoderinae (von Horn, 1881) Jeannel
18 Subfamily Ditominae
19 Subfamily Anisodactylinae
20 Subfamily Harpalinae Bonelli, 1810
21 Subfamily Stenolophinae
22 Subfamily Amblystominae
23 Subfamily Pterostichinae
25 Subfamily Zabrinae
26 Subfamily Callistinae Habe & Kosuge, 1967
27 Subfamily Oodinae
28 Subfamily Licininae Bonelli, 1810
29 Subfamily Badistrinae
30 Subfamily Panagaeinae
31 Subfamily Colliurinae
32 Subfamily Aephnidiinae
33 Subfamily Lebiinae
34 Subfamily Demetriinae
35 Subfamily Callidinae
36 Subfamily Cymindinae
37 Subfamily Dromiinae De Haan, 1833 [in De Haan, 1833-1850]
38 Subfamily Zuphiinae
39 Subfamily Dryptinae
40 Subfamily Mormolycinae
41 Subfamily Brachininae Bonelli, 1810
42 Subfamily Migadopinae Chaudoir, 1861
43 Subfamily Paussinae (Latreille, 1806)