Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade neomura Cavalier-Smith, 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade discaria Cavalier-Smith, 2021
                 Clade dorsates Cavalier-Smith, 2021
                   Clade podiates Cavalier-Smith, 2012
                     Clade Amorphea Adl et al. 2012 [Simpson, Medina & Adl 2020]
                       Clade Obazoa Brown, Sharpe, Silberman, Heiss, Lang, Simpson & Roger 2013
                         Clade Opisthokonta Cavalier-Smith 1987 [Simpson & Medina 2020] - animal-fungal clade
                           Superkingdom Holomycota Liu, Steenkamp, Brinkmann, Forget, Philippe & Lang, 2009
                             Kingdom Fungi R.T.Moore (1980) - fungi
                               Subkingdom Dikarya Hibbett, T.Y. James & Vilgalys (2007)
                                 Phylum Ascomycota Caval.-Sm. (1998) - sac fungi
                                   Subphylum Pezizomycotina O.E. Erikss. & Winka (1997)
                                     Class Eurotiomycetes O.E. Erikss. & Winka (1997)
                                       Subclass Chaetothyriomycetidae Doweld (2001)
                                         Order Pyrenulales Fink ex D.Hawksw. & O.E. Erikss. (1986) [monotypic]
                                           Family Pyrenulaceaeᵀ Rabenh. (1870)
                                             01 Genus Acrocordiella O.E. Erikss. (1982)
                                             02 Genus Anthracothecium Hampe ex A.Massal. (1860)
                                             03 Genus Blastodesmia A.Massal. (1852)
                                             04 Genus Clypeopyrenis Aptroot (1991)
                                             05 Genus Distopyrenis Aptroot (1991)
                                             06 Genus Granulopyrenis Aptroot (1991)
                                             07 Genus Lithothelium Müll.Arg. (1885)
                                             08 Genus Mazaediothecium Aptroot (1991)
                                             09 Genus Pyrenographa Aptroot (1991)
                                             10 Genus Pyrenowilmsia R.C. Harris & Aptroot (1991)
                                             11 Genus Pyrenulaᵀ Ach. (1814), nom. cons.
                                             12 Genus Pyrgillus Nyl. (1858)
                                             13 Genus Sulcopyrenula H. Harada (1999)