[organisms] see Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
   Pisces
     Class Osteichthyes Huxley, 1880 - bony fishes and tetrapods see Clade Osteichthyes Huxley 1880 [Moore & Near 2020]
       Order Gadiformes Goodrich, 1909 - cods and hakes
         Family Merlucciidae Rafinesque, 1815 - merlucciid hakes
           Genus Macruronus Günther, 1873
             Subgenus Macruronus (Cynogadus) Howes, 1991
               Macruronus (Cynogadus) brachycolus (Holt & Byrne, 1906) see Lyconus brachycolus Holt & Byrne, 1906