Natura - nature
   actualia - actual entities
     Mundus Plinius - physical world
       naturalia - natural bodies
         Clade Biota Wagner 2004 [Wiemann, de Queiroz, Rowe, Planavsky, Anderson, Gogarten, Turner & Gauthier 2020]
           Clade neomura Cavalier-Smith, 1987
             Domain Eukaryota Chatton, 1925 - eukaryotes
               Clade discaria Cavalier-Smith, 2021
                 Clade natates Cavalier-Smith, 2021
                   Clade photaria Cavalier-Smith, 2021
                     Clade eucorta Cavalier-Smith, 2021
                       Superkingdom Corticata Lankester, 1878 - corticates
                         Kingdom Chromista Caval.-Sm. (1981) - chromists
                           Subkingdom Harosa Cavalier-Smith, 2010
                             Clade Sar Adl et al. 2012 [Simpson & Dunthorn 2020]
                               Infrakingdom Rhizaria Cavalier-Smith, 2002
                                 Phylum Cercozoa Cavalier-Smith, 1998
                                   Subphylum Endomyxa T. Cavalier-Smith, 2002
                                     Class Phytomyxea Engler & Prantl, 1897
                                       Order Plasmodiophorida Cook, 1928
                                         Family Plasmodiophoridaeᵀ Zopf, 1884
                                           Genus Ligniera Maire & Tison, 1911
                                             Ligniera pilorum Fron & Gaillat
                                             Ligniera verrucosa Maire & A. Tison